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Selected publications
(last five years only)
(last five years only)
Underlined authors are Sharma group members.
For complete bibliography, please download CVs on the personnel page.
In preparation
85. *Crawford, A.R., Gainett, G., *So, C., Sharma, P.P. (in prep.) Eggs to long-legs: Embryonic development and staging of the harvestman Phalangium opilio (Arachnida, Opiliones). (*undergraduate co-author)
84. Setton, E.V.W., Sharma, P.P. (in prep.) Morphogenesis of spider spinnerets.
83. Setton, E.V.W., Sharma, P.P. (in prep.) Systemic effects of inhibiting Wnt signaling in the spider Parasteatoda tepidariorum.
82. Santibáñez-López, C.E., Nolan, E.D., Sharma, P.P. (in prep.) Morphogenesis of the scorpion pectine.
81. Nolan, E.D., Santibáñez-López, C.E., Gainett, G., Ontano, A.Z., Setton, E.V.W., Sharma, P.P. (in prep.) Developmental data as a phylogenetic character.
Peer reviewed
80. Cano-Sánchez, E., Sharma, P.P., López-González, P.J. (in review) Postembryonic development of Nymphon australe Hodgson, 1902 (Pycnogonida, Nymphonidae) from Antarctica. Polar Biology.
79. Aharon, S., Ballesteros, J.A., Crawford, A.C., Friske, K., Guilherme, G., Langford, B., Santibáñez-López, C.E., Ya'aran, S., Gavish-Regev, E. & Sharma, P.P. (2019) The anatomy of an unstable node: A Levantine relict precipitates phylogenomic dissolution of higher-level relationships of the armored harvestmen (Arachnida: Opiliones: Laniatores). Invertebrate Systematics (doi: https://doi.org/10.1071/IS18057)
78. Santibáñez-López, C.E., González-Santillán, E., Monod, L., Sharma, P.P. (2019) Phylogenomics facilitates stable scorpion systematics: Reassessing the relationships of Vaejovidae and a new high-level classification of Scorpiones (Arachnida). Molecular Phylogenetics and Evolution 135:22-30. [PDF]
77. Setton, E.V.W., Hendrixson, B.E. & Sharma, P.P. (in press) Embryogenesis in a Colorado population of Aphonopelma hentzi (Girard, 1852), a promising system for the study of development (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology.
76. Gainett, G., Sharma, P.P., Fernandes, N., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2019) Evolution of a sensory cluster on the legs of Opiliones (Arachnida) informs multi-level phylogenetic relationships. Zoological Journal of the Linnean Society (doi: doi/10.1093/zoolinnean/zlz016/5437571) [PDF]
75. Ballesteros, J.A. & Sharma, P.P. (2019) A critical appraisal of the placement of Xiphosura (Chelicerata) with account of known sources of phylogenetic error. Systematic Biology (doi: 10.1093/sysbio/syz011) [PDF]
74. Sharma, P.P. (2019) Integrating morphology and phylogenomics supports a terrestrial origin of insect flight. Proceedings of the National Academy of Sciences of the USA 116:2796-2798. [PDF]
73. *Schmidt, S.M., Buenavente, P.A.C., Blatchley, D.D., Diesmos, A.C., Diesmos, M.L., General, D.E.M., Mohagan, A.B., Mohagan, D.J., Clouse, R.M. & Sharma, P.P. (2019) A new species of Tithaeidae (Arachnida: Opiliones: Laniatores) from Mindanao reveals contemporaneous colonization of the Philippines by Sunda Shelf opiliofauna. Invertebrate Systematics 33:237-251. [PDF]
72. *March, L.E., *Smaby, R.M., *Setton, E.V.W. & Sharma, P.P. (2018) The evolution of selector gene function: Expression dynamics and regulatory interactions of tiptop/teashirt across Arthropoda. Evolution & Development 20:219-232. [PDF] (*undergraduate co-author)
71. Santibañez López, C.E., Kriebel, R., Ballesteros, J.A., Rush, N., Witter, Z., Williams, J., Janies, D.A. & Sharma, P.P. (2018) Integration of phylogenomics and molecular modeling reveals lineage-specific diversification of toxins in scorpions. PeerJ 5902:1-13. [PDF]
70. Zhang, C., Zhang F. & Sharma, P.P. (2018) Two new species of Petrobunus Sharma & Giribet, 2011 from China (Opiliones: Laniatores: Petrobunidae). Zootaxa 4524:51-64. [PDF]
69. Leite, D.J., Baudouin-Gonzalez, L., Iwasaki-Yokozawa, S., Lozano-Fernandez, J., Turetzek, N., Akiyama-Oda, Y., Pisani, D., Oda, H., Sharma, P.P. & McGregor, A.P. (2018) Homeobox gene duplication and divergence in arachnids. Molecular Biology and Evolution 35:2240-2253. [PDF]
68. Sharma, P.P. (2018) Chelicerates. Current Biology 28:R774-778. [PDF]
67. Oberski, J.T., Sharma, P.P., Jay, K.R., Coblens, M., Lemon, K.A., Johnson, J.E. & Boyer, S.L. (2018) A dated molecular phylogeny of mite harvestmen (Arachnida: Opiliones: Cyphophthalmi) elucidates ancient diversification dynamics in the Australian Wet Tropics. Molecular Phylogenetics and Evolution (doi: 10.1016/j.ympev.2018.06.029) [PDF].
66. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) The sensory equipment of a sandokanid: an extreme case of tarsal reduction in a harvestman (Arachnida, Opiliones). Journal of Morphology (doi: 10.1002/jmor.20843). [PDF]
65. Di, Z., Edgecombe, G.D., Sharma, P.P. (2018) Homeosis in a scorpion supports a telopodal origin of pectines and book lungs. BMC Evolutionary Biology 18:73. [PDF]
64. Santibáñez-López, C.E., Ontano, A.Z., Harvey, M.S., & Sharma, P.P. (2018) Transcriptomic analysis of pseudoscorpion venom reveals a unique cocktail dominated by enzymes and protease inhibitors. Toxins 10:207. [PDF]
63. *Setton, E.V.W., Sharma, P.P. (2018) Cooption of an appendage patterning gene cassette in the head segmentation of arachnids. Proceedings of the National Academy of Sciences of the USA 115:E3491-E3500. [PDF] (*undergraduate co-author)
62. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) Putative adhesive setae on the walking legs of the Paleotropical harvestman Metibalonius sp. (Arachnida: Opiliones: Podoctidae). Journal of Arachnology 46:62-68. [PDF]
61. Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. & Giribet, G. (2018) A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122:37-45. [PDF]
60. Garb, J.E., Sharma, P.P. & Ayoub, N.A. (2018) Recent progress and prospects for advancing arachnid genomics. Current Opinions in Insect Science 25:51-57. [PDF]
59. Lewis, S.H., Quarles, K.A., Yang, Y., Tanguy, M., Frézal, L., Smith, S., Sharma, P.P, Cordaux, R., Gilbert, C., Giraud, I., Collins, D.H., Zamore, P.D., Miska, E.A., Sarkies, P. & Jiggins, F.M. (2018) Pan-arthropod analysis reveals somatic piRNAs as an ancestral defence against transposable elements. Nature Ecology and Evolution 2:174-181. [PDF]
58. Daniels, S.R., Dreyer, M. & Sharma, P.P. (2017) Contrasting the population genetic structure of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsis) in forest fragments along the south eastern Cape, South Africa. Invertebrate Systematics 31:781-796. [PDF]
57. Sharma, P.P. (2017) Chelicerates and the conquest of land: A view of arachnid origins through an evodevo spyglass. Integrative and Comparative Biology 57, 510-522. [PDF]
56. *Schwager, E.E., *Sharma, P.P., *Clarke, T., *Leite, D.J., Wierschin, T., Pechmann M., Akiyama-Oda, Y., Esposito, L., Arensburger, P., Bechsgaard, J., Bilde, T., Buffry, A., Chao, H., Dinh, H., Doddapaneni, H., Dugan, S., Eibner C., Extavour, C.G., Funch, P., Garb, J., Gonzalez, V.L., Griffiths-Jones, S., Han, Y., Hayashi, C., Hilbrant, M., Hughes, D.S.T., Janssen, R., Lee, S.L., Maeso, I., Murali, S.C., Muzny, D.M., da Fonseca, R.N., Qu, J., Ronshaugen, M., Schomburg, C., Schönauer, A., Stollewerk, A., Torres-Oliva, M., Turetzek, N., Vanthournout, B., Werren, J., Wolff, C., Worley, K.C., Gibbs, R.A., Coddington, J., Oda, H., Stanke, M., Ayoub, N.A., Damen, W.G.M., Prpic, N.-M., Flot, J.-F., Posnien, N., Richards, S., & McGregor, A.P. (2017) The house spider genome reveals a whole genome duplication during arachnid evolution. BMC Biology 15:62. [PDF] (*equal author contribution)
55. *Setton, E.V.W., *March, L.E., *Nolan, E.D., Jones, T.E., Cho, H., Wheeler, W.C., Extavour, C.G. & Sharma, P.P. (2017) Expression and function of spineless orthologs correlate with distal deutocerebral appendage morphology in Arthropoda. Developmental Biology 430:224-236. (*undergraduate co-author) [PDF]
54. Clouse, R.M., Branstetter, M.G., Buenavente, P.A.C., Crowley, L.M., Czekanski-Moir, J., General, D.E.M., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Sharma, P.P. & Wheeler, W.C. (2017) First molecular phylogeny of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44:2660-2672. [PDF] [Featured on the journal cover]
53. Sharma P.P., Oberski, J.T., Santiago, M.A., Ricardo Kriebel, *Savana M. Lipps, Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). Molecular Phylogenetics and Evolution. (*undergraduate co-author) (doi: 10.1016/j.ympev.2017.03.026) [PDF]
52. *Santibañez-López, C., *Kriebel, R. & Sharma, P.P. (2017) eadem figura manet: Measuring morphological convergence in diplocentrid scorpions (Arachnida: Scorpiones: Diplocentridae) under a multilocus phylogenetic framework. Invertebrate Systematics. Invertebrate Systematics 31:233-248. [PDF] (*equal author contribution) [Featured on the journal cover]
51. Fernández, R., Sharma P.P., Tourinho, A.L. & Giribet, G. (2017) The Opiliones Tree of Life: shedding light on harvestmen relationships through transcriptomics. Proceedings of the Royal Society of London B: Biological Sciences 284:20162340. [PDF]
50. Sharma P.P., *Santiago, M.A., Kriebel, R., *Lipps, S.M., Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) reveals the disutility of subfamily nomenclature and traditional character systems in armored harvestman systematics. Molecular Phylogenetics and Evolution 106:164-173. [PDF] (*undergraduate co-author)
49. *Jay, K.R., Popkin-Hall, Z.R., *Coblens, M.J., *Oberski, J.T., Sharma, P.P. & Boyer, S.L. (2016) New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot. Zookeys 586:37-93. [PDF] (*undergraduate co-author)
48. Iyer, H., Issigonis, M., Sharma, P.P., Extavour, C.G. & Newmark, P.A. (2016) A novel pre-meiotic function for boule in the planarian Schmidtea mediterranea. Proceedings of the National Academy of Sciences of the USA 113:E3509-E3518. (doi: 10.1073/pnas.1521341113) [PDF]
47. Sharma, P.P., Clouse, R.M. & Wheeler, W.C. (2016) Hennig's semaphoront concept and the use of ontogenetic stages in phylogenetic reconstruction. Cladistics. (doi: 10.1111/cla.12156) [PDF]
46. Daniels, S.R., Kunaka, C., Klaus, S. & Sharma, P.P. (2016) Unmasking alpha diversity, cladogenesis and biogeographical patterning in an ancient panarthropod lineage (Onychophora: Peripatopsidae: Opisthopatus cinctipes) with the description of five novel species. Cladistics. (doi: 10.1111/cla.12154) [PDF]
45. Giribet, G., Boyer, S.L., Baker, C.M., Fernández, R., Sharma P.P., de Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. & Griswold, C.E. (2016) A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178:523-545. [PDF]
44. Pérez-González, A., Sharma, P.P. & Proud, D.N. (2016) Morphological tricks and blessed genitalia: rectifying the familial placement of Fijicolana tuberculata (Opiliones: Laniatores: Zalmoxidae). Zootaxa 4061:253-260. [PDF]
43. Clouse, R.M., Sharma, P.P., *Stuart, J.C., Davis, L.R., Giribet, G., Sarah, S.L. & Wheeler, W.C. (2016) Phylogeography of the harvestman Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox. Organisms, Diversity and Evolution 16:167-184. [PDF] (*undergraduate co-author)
42. Sharma, P.P., *Santiago, M.A., González-Santillán, E., Monod, L. & Wheeler, W.C. (2015) Evidence of duplicated Hox genes in the most recent common ancestor of extant scorpions. Evolution and Development 17:347-355. [PDF] (*undergraduate co-author)
41. Clouse, R.M., Janda, M., *Blanchard, B., Sharma, P.P., Hoffmann, B.D., Andersen, A.N., Czekanski-Moir, J.E., Krushelnycky, P. Rabeling, C., Wilson, E.O., Economo, E.P., Sarnat, E.M., General, D.M., Alpert, G.D. & Wheeler, W.C. (2015) Molecular phylogeny of an Indo-Pacific carpenter ant (Hymenoptera: Formicidae, Camponotus) reveals waves of dispersal and colonization from diverse sources. Cladistics 31:424-437. [PDF] (*undergraduate co-author)
40. Schwager, E.E., Schönauer, A., Leite, D.J., Sharma, P.P. & McGregor, A.P. (2015) Chelicerata. In: Wanninger, A. (editor) Evolutionary Developmental Biology of Invertebrates, Vol. 3: Ecdysozoa I: Non-Tetraconata. Heidelberg: Springer. [PDF]
39. Sharma, P.P., Tarazona, O.A., Lopez, D.H., Schwager, E.E., Cohn, M.J., Wheeler, W.C. & Extavour, C.G. (2015) A conserved genetic mechanism differentiates gnathal and locomotory appendages in insects and arachnids. Proceedings of the Royal Society of London B: Biological Sciences 282: 20150698. [PDF]
38. Sharma, P.P., Fernández, R., Esposito, L., González-Santillán, E. & Monod, L. (2015). Phylogenomic resolution of scorpions reveals multilevel discordance with morphological phylogenetic signal. Proceedings of the Royal Society of London B: Biological Sciences 282: 20142953. [PDF]
37. Giribet, G. & Sharma, P.P. (2015) Evolutionary biology of harvestmen (Arachnida, Opiliones). Annual Reviews of Entomology 60:157-175. [PDF]
36. Sharma, P.P., Kaluziak, S.T., Pérez-Porro, A.R., González, V.L., Hormiga, G., Wheeler, W.C. & Giribet, G. (2014) Phylogenomic interrogation of Arachnida reveals systemic conflicts in phylogenetic signal. Molecular Biology and Evolution 31:2963-2984. [PDF] [Featured in the MBE Citation Classics of 2018, the top 1% of highly cited works]
35. Sharma, P.P. & Wheeler, W.C. (2014) Cross-bracing uncalibrated nodes in molecular dating improves congruence of fossil and molecular age estimates. Frontiers in Zoology 11:57. [PDF]
34. Sharma, P.P., Schwager, E.E., Extavour, C.G. & Wheeler, W.C. (2014) Hox gene duplications correlate with posterior heteronomy in scorpions. Proceedings of the Royal Society of London B: Biological Sciences 281:20140661. [Featured in AMNH News & Blogs] [PDF]
33. Sharma, P.P. & Giribet G. (2014) A revised, dated phylogeny of the arachnid order Opiliones. Frontiers in Genetics 5:255. [PDF]
32. Fernandéz, R., Vahtera, V., Laumer, C.E., Libro, S., Kaluziak, S.T., Sharma, P.P., Pérez-Porro, A.R., Edgecombe, G.D. & Giribet, G. (2014) Evaluating topological conflict in centipede phylogeny using transcriptomic data sets. Molecular Biology and Evolution 31:1500-1513. [PDF]
31. *Garwood, R., *Sharma, P.P., Dunlop, J.A & Giribet, G. (2014) A Palaeozoic stem-group to mite harvestman revealed through integration of phylogenetics and development. (*equal author contribution). Current Biology 24:1017-1023. [Featured in: National Geographic's Weird and Wild; Discovery News; LiveScience; Wired; PBS Newshour; Huffington Post; International Business Times; Buzzfeed] [PDF]
30. Bieler, R., Mikkelsen, P.M., Collins, T.M., Glover, E.A., González, V.L., Graf, D.L., Harper, E.M., Healy, J., Kawauchi, G.Y., Sharma, P.P., Staubach, S., Strong, E.E., Taylor, J.D., Temkin, I., Zardus, J.D., Clark, S., Guzmán, A., McIntyre, E., Sharp, P. & Giribet, G. (2014) Investigating the Bivalve Tree of Life—an exemplar-based approach combining molecular and novel morphological characters. Invertebrate Systematics 28:32-115. [Featured on the journal cover] [compressed PDF]
29. Sharma, P.P., Gupta, T., Schwager, E.E., Wheeler, W.C. & Extavour, C.G. (2014) Subdivision of arthropod cap-n-collar expression domains is restricted to Mandibulata. EvoDevo 5:3. [PDF]
28. *Gainett, G., Sharma, P.P., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2014) Walk it off: Predictive power of appendicular characters toward inference of higher-level relationships in Laniatores (Arachnida: Opiliones). Cladistics 30:120-138. [PDF] (*undergraduate co-author)
For complete bibliography, please download CVs on the personnel page.
In preparation
85. *Crawford, A.R., Gainett, G., *So, C., Sharma, P.P. (in prep.) Eggs to long-legs: Embryonic development and staging of the harvestman Phalangium opilio (Arachnida, Opiliones). (*undergraduate co-author)
84. Setton, E.V.W., Sharma, P.P. (in prep.) Morphogenesis of spider spinnerets.
83. Setton, E.V.W., Sharma, P.P. (in prep.) Systemic effects of inhibiting Wnt signaling in the spider Parasteatoda tepidariorum.
82. Santibáñez-López, C.E., Nolan, E.D., Sharma, P.P. (in prep.) Morphogenesis of the scorpion pectine.
81. Nolan, E.D., Santibáñez-López, C.E., Gainett, G., Ontano, A.Z., Setton, E.V.W., Sharma, P.P. (in prep.) Developmental data as a phylogenetic character.
Peer reviewed
80. Cano-Sánchez, E., Sharma, P.P., López-González, P.J. (in review) Postembryonic development of Nymphon australe Hodgson, 1902 (Pycnogonida, Nymphonidae) from Antarctica. Polar Biology.
79. Aharon, S., Ballesteros, J.A., Crawford, A.C., Friske, K., Guilherme, G., Langford, B., Santibáñez-López, C.E., Ya'aran, S., Gavish-Regev, E. & Sharma, P.P. (2019) The anatomy of an unstable node: A Levantine relict precipitates phylogenomic dissolution of higher-level relationships of the armored harvestmen (Arachnida: Opiliones: Laniatores). Invertebrate Systematics (doi: https://doi.org/10.1071/IS18057)
78. Santibáñez-López, C.E., González-Santillán, E., Monod, L., Sharma, P.P. (2019) Phylogenomics facilitates stable scorpion systematics: Reassessing the relationships of Vaejovidae and a new high-level classification of Scorpiones (Arachnida). Molecular Phylogenetics and Evolution 135:22-30. [PDF]
77. Setton, E.V.W., Hendrixson, B.E. & Sharma, P.P. (in press) Embryogenesis in a Colorado population of Aphonopelma hentzi (Girard, 1852), a promising system for the study of development (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology.
76. Gainett, G., Sharma, P.P., Fernandes, N., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2019) Evolution of a sensory cluster on the legs of Opiliones (Arachnida) informs multi-level phylogenetic relationships. Zoological Journal of the Linnean Society (doi: doi/10.1093/zoolinnean/zlz016/5437571) [PDF]
75. Ballesteros, J.A. & Sharma, P.P. (2019) A critical appraisal of the placement of Xiphosura (Chelicerata) with account of known sources of phylogenetic error. Systematic Biology (doi: 10.1093/sysbio/syz011) [PDF]
74. Sharma, P.P. (2019) Integrating morphology and phylogenomics supports a terrestrial origin of insect flight. Proceedings of the National Academy of Sciences of the USA 116:2796-2798. [PDF]
73. *Schmidt, S.M., Buenavente, P.A.C., Blatchley, D.D., Diesmos, A.C., Diesmos, M.L., General, D.E.M., Mohagan, A.B., Mohagan, D.J., Clouse, R.M. & Sharma, P.P. (2019) A new species of Tithaeidae (Arachnida: Opiliones: Laniatores) from Mindanao reveals contemporaneous colonization of the Philippines by Sunda Shelf opiliofauna. Invertebrate Systematics 33:237-251. [PDF]
72. *March, L.E., *Smaby, R.M., *Setton, E.V.W. & Sharma, P.P. (2018) The evolution of selector gene function: Expression dynamics and regulatory interactions of tiptop/teashirt across Arthropoda. Evolution & Development 20:219-232. [PDF] (*undergraduate co-author)
71. Santibañez López, C.E., Kriebel, R., Ballesteros, J.A., Rush, N., Witter, Z., Williams, J., Janies, D.A. & Sharma, P.P. (2018) Integration of phylogenomics and molecular modeling reveals lineage-specific diversification of toxins in scorpions. PeerJ 5902:1-13. [PDF]
70. Zhang, C., Zhang F. & Sharma, P.P. (2018) Two new species of Petrobunus Sharma & Giribet, 2011 from China (Opiliones: Laniatores: Petrobunidae). Zootaxa 4524:51-64. [PDF]
69. Leite, D.J., Baudouin-Gonzalez, L., Iwasaki-Yokozawa, S., Lozano-Fernandez, J., Turetzek, N., Akiyama-Oda, Y., Pisani, D., Oda, H., Sharma, P.P. & McGregor, A.P. (2018) Homeobox gene duplication and divergence in arachnids. Molecular Biology and Evolution 35:2240-2253. [PDF]
68. Sharma, P.P. (2018) Chelicerates. Current Biology 28:R774-778. [PDF]
67. Oberski, J.T., Sharma, P.P., Jay, K.R., Coblens, M., Lemon, K.A., Johnson, J.E. & Boyer, S.L. (2018) A dated molecular phylogeny of mite harvestmen (Arachnida: Opiliones: Cyphophthalmi) elucidates ancient diversification dynamics in the Australian Wet Tropics. Molecular Phylogenetics and Evolution (doi: 10.1016/j.ympev.2018.06.029) [PDF].
66. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) The sensory equipment of a sandokanid: an extreme case of tarsal reduction in a harvestman (Arachnida, Opiliones). Journal of Morphology (doi: 10.1002/jmor.20843). [PDF]
65. Di, Z., Edgecombe, G.D., Sharma, P.P. (2018) Homeosis in a scorpion supports a telopodal origin of pectines and book lungs. BMC Evolutionary Biology 18:73. [PDF]
64. Santibáñez-López, C.E., Ontano, A.Z., Harvey, M.S., & Sharma, P.P. (2018) Transcriptomic analysis of pseudoscorpion venom reveals a unique cocktail dominated by enzymes and protease inhibitors. Toxins 10:207. [PDF]
63. *Setton, E.V.W., Sharma, P.P. (2018) Cooption of an appendage patterning gene cassette in the head segmentation of arachnids. Proceedings of the National Academy of Sciences of the USA 115:E3491-E3500. [PDF] (*undergraduate co-author)
62. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) Putative adhesive setae on the walking legs of the Paleotropical harvestman Metibalonius sp. (Arachnida: Opiliones: Podoctidae). Journal of Arachnology 46:62-68. [PDF]
61. Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. & Giribet, G. (2018) A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122:37-45. [PDF]
60. Garb, J.E., Sharma, P.P. & Ayoub, N.A. (2018) Recent progress and prospects for advancing arachnid genomics. Current Opinions in Insect Science 25:51-57. [PDF]
59. Lewis, S.H., Quarles, K.A., Yang, Y., Tanguy, M., Frézal, L., Smith, S., Sharma, P.P, Cordaux, R., Gilbert, C., Giraud, I., Collins, D.H., Zamore, P.D., Miska, E.A., Sarkies, P. & Jiggins, F.M. (2018) Pan-arthropod analysis reveals somatic piRNAs as an ancestral defence against transposable elements. Nature Ecology and Evolution 2:174-181. [PDF]
58. Daniels, S.R., Dreyer, M. & Sharma, P.P. (2017) Contrasting the population genetic structure of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsis) in forest fragments along the south eastern Cape, South Africa. Invertebrate Systematics 31:781-796. [PDF]
57. Sharma, P.P. (2017) Chelicerates and the conquest of land: A view of arachnid origins through an evodevo spyglass. Integrative and Comparative Biology 57, 510-522. [PDF]
56. *Schwager, E.E., *Sharma, P.P., *Clarke, T., *Leite, D.J., Wierschin, T., Pechmann M., Akiyama-Oda, Y., Esposito, L., Arensburger, P., Bechsgaard, J., Bilde, T., Buffry, A., Chao, H., Dinh, H., Doddapaneni, H., Dugan, S., Eibner C., Extavour, C.G., Funch, P., Garb, J., Gonzalez, V.L., Griffiths-Jones, S., Han, Y., Hayashi, C., Hilbrant, M., Hughes, D.S.T., Janssen, R., Lee, S.L., Maeso, I., Murali, S.C., Muzny, D.M., da Fonseca, R.N., Qu, J., Ronshaugen, M., Schomburg, C., Schönauer, A., Stollewerk, A., Torres-Oliva, M., Turetzek, N., Vanthournout, B., Werren, J., Wolff, C., Worley, K.C., Gibbs, R.A., Coddington, J., Oda, H., Stanke, M., Ayoub, N.A., Damen, W.G.M., Prpic, N.-M., Flot, J.-F., Posnien, N., Richards, S., & McGregor, A.P. (2017) The house spider genome reveals a whole genome duplication during arachnid evolution. BMC Biology 15:62. [PDF] (*equal author contribution)
55. *Setton, E.V.W., *March, L.E., *Nolan, E.D., Jones, T.E., Cho, H., Wheeler, W.C., Extavour, C.G. & Sharma, P.P. (2017) Expression and function of spineless orthologs correlate with distal deutocerebral appendage morphology in Arthropoda. Developmental Biology 430:224-236. (*undergraduate co-author) [PDF]
54. Clouse, R.M., Branstetter, M.G., Buenavente, P.A.C., Crowley, L.M., Czekanski-Moir, J., General, D.E.M., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Sharma, P.P. & Wheeler, W.C. (2017) First molecular phylogeny of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44:2660-2672. [PDF] [Featured on the journal cover]
53. Sharma P.P., Oberski, J.T., Santiago, M.A., Ricardo Kriebel, *Savana M. Lipps, Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). Molecular Phylogenetics and Evolution. (*undergraduate co-author) (doi: 10.1016/j.ympev.2017.03.026) [PDF]
52. *Santibañez-López, C., *Kriebel, R. & Sharma, P.P. (2017) eadem figura manet: Measuring morphological convergence in diplocentrid scorpions (Arachnida: Scorpiones: Diplocentridae) under a multilocus phylogenetic framework. Invertebrate Systematics. Invertebrate Systematics 31:233-248. [PDF] (*equal author contribution) [Featured on the journal cover]
51. Fernández, R., Sharma P.P., Tourinho, A.L. & Giribet, G. (2017) The Opiliones Tree of Life: shedding light on harvestmen relationships through transcriptomics. Proceedings of the Royal Society of London B: Biological Sciences 284:20162340. [PDF]
50. Sharma P.P., *Santiago, M.A., Kriebel, R., *Lipps, S.M., Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) reveals the disutility of subfamily nomenclature and traditional character systems in armored harvestman systematics. Molecular Phylogenetics and Evolution 106:164-173. [PDF] (*undergraduate co-author)
49. *Jay, K.R., Popkin-Hall, Z.R., *Coblens, M.J., *Oberski, J.T., Sharma, P.P. & Boyer, S.L. (2016) New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot. Zookeys 586:37-93. [PDF] (*undergraduate co-author)
48. Iyer, H., Issigonis, M., Sharma, P.P., Extavour, C.G. & Newmark, P.A. (2016) A novel pre-meiotic function for boule in the planarian Schmidtea mediterranea. Proceedings of the National Academy of Sciences of the USA 113:E3509-E3518. (doi: 10.1073/pnas.1521341113) [PDF]
47. Sharma, P.P., Clouse, R.M. & Wheeler, W.C. (2016) Hennig's semaphoront concept and the use of ontogenetic stages in phylogenetic reconstruction. Cladistics. (doi: 10.1111/cla.12156) [PDF]
46. Daniels, S.R., Kunaka, C., Klaus, S. & Sharma, P.P. (2016) Unmasking alpha diversity, cladogenesis and biogeographical patterning in an ancient panarthropod lineage (Onychophora: Peripatopsidae: Opisthopatus cinctipes) with the description of five novel species. Cladistics. (doi: 10.1111/cla.12154) [PDF]
45. Giribet, G., Boyer, S.L., Baker, C.M., Fernández, R., Sharma P.P., de Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. & Griswold, C.E. (2016) A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178:523-545. [PDF]
44. Pérez-González, A., Sharma, P.P. & Proud, D.N. (2016) Morphological tricks and blessed genitalia: rectifying the familial placement of Fijicolana tuberculata (Opiliones: Laniatores: Zalmoxidae). Zootaxa 4061:253-260. [PDF]
43. Clouse, R.M., Sharma, P.P., *Stuart, J.C., Davis, L.R., Giribet, G., Sarah, S.L. & Wheeler, W.C. (2016) Phylogeography of the harvestman Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox. Organisms, Diversity and Evolution 16:167-184. [PDF] (*undergraduate co-author)
42. Sharma, P.P., *Santiago, M.A., González-Santillán, E., Monod, L. & Wheeler, W.C. (2015) Evidence of duplicated Hox genes in the most recent common ancestor of extant scorpions. Evolution and Development 17:347-355. [PDF] (*undergraduate co-author)
41. Clouse, R.M., Janda, M., *Blanchard, B., Sharma, P.P., Hoffmann, B.D., Andersen, A.N., Czekanski-Moir, J.E., Krushelnycky, P. Rabeling, C., Wilson, E.O., Economo, E.P., Sarnat, E.M., General, D.M., Alpert, G.D. & Wheeler, W.C. (2015) Molecular phylogeny of an Indo-Pacific carpenter ant (Hymenoptera: Formicidae, Camponotus) reveals waves of dispersal and colonization from diverse sources. Cladistics 31:424-437. [PDF] (*undergraduate co-author)
40. Schwager, E.E., Schönauer, A., Leite, D.J., Sharma, P.P. & McGregor, A.P. (2015) Chelicerata. In: Wanninger, A. (editor) Evolutionary Developmental Biology of Invertebrates, Vol. 3: Ecdysozoa I: Non-Tetraconata. Heidelberg: Springer. [PDF]
39. Sharma, P.P., Tarazona, O.A., Lopez, D.H., Schwager, E.E., Cohn, M.J., Wheeler, W.C. & Extavour, C.G. (2015) A conserved genetic mechanism differentiates gnathal and locomotory appendages in insects and arachnids. Proceedings of the Royal Society of London B: Biological Sciences 282: 20150698. [PDF]
38. Sharma, P.P., Fernández, R., Esposito, L., González-Santillán, E. & Monod, L. (2015). Phylogenomic resolution of scorpions reveals multilevel discordance with morphological phylogenetic signal. Proceedings of the Royal Society of London B: Biological Sciences 282: 20142953. [PDF]
37. Giribet, G. & Sharma, P.P. (2015) Evolutionary biology of harvestmen (Arachnida, Opiliones). Annual Reviews of Entomology 60:157-175. [PDF]
36. Sharma, P.P., Kaluziak, S.T., Pérez-Porro, A.R., González, V.L., Hormiga, G., Wheeler, W.C. & Giribet, G. (2014) Phylogenomic interrogation of Arachnida reveals systemic conflicts in phylogenetic signal. Molecular Biology and Evolution 31:2963-2984. [PDF] [Featured in the MBE Citation Classics of 2018, the top 1% of highly cited works]
35. Sharma, P.P. & Wheeler, W.C. (2014) Cross-bracing uncalibrated nodes in molecular dating improves congruence of fossil and molecular age estimates. Frontiers in Zoology 11:57. [PDF]
34. Sharma, P.P., Schwager, E.E., Extavour, C.G. & Wheeler, W.C. (2014) Hox gene duplications correlate with posterior heteronomy in scorpions. Proceedings of the Royal Society of London B: Biological Sciences 281:20140661. [Featured in AMNH News & Blogs] [PDF]
33. Sharma, P.P. & Giribet G. (2014) A revised, dated phylogeny of the arachnid order Opiliones. Frontiers in Genetics 5:255. [PDF]
32. Fernandéz, R., Vahtera, V., Laumer, C.E., Libro, S., Kaluziak, S.T., Sharma, P.P., Pérez-Porro, A.R., Edgecombe, G.D. & Giribet, G. (2014) Evaluating topological conflict in centipede phylogeny using transcriptomic data sets. Molecular Biology and Evolution 31:1500-1513. [PDF]
31. *Garwood, R., *Sharma, P.P., Dunlop, J.A & Giribet, G. (2014) A Palaeozoic stem-group to mite harvestman revealed through integration of phylogenetics and development. (*equal author contribution). Current Biology 24:1017-1023. [Featured in: National Geographic's Weird and Wild; Discovery News; LiveScience; Wired; PBS Newshour; Huffington Post; International Business Times; Buzzfeed] [PDF]
30. Bieler, R., Mikkelsen, P.M., Collins, T.M., Glover, E.A., González, V.L., Graf, D.L., Harper, E.M., Healy, J., Kawauchi, G.Y., Sharma, P.P., Staubach, S., Strong, E.E., Taylor, J.D., Temkin, I., Zardus, J.D., Clark, S., Guzmán, A., McIntyre, E., Sharp, P. & Giribet, G. (2014) Investigating the Bivalve Tree of Life—an exemplar-based approach combining molecular and novel morphological characters. Invertebrate Systematics 28:32-115. [Featured on the journal cover] [compressed PDF]
29. Sharma, P.P., Gupta, T., Schwager, E.E., Wheeler, W.C. & Extavour, C.G. (2014) Subdivision of arthropod cap-n-collar expression domains is restricted to Mandibulata. EvoDevo 5:3. [PDF]
28. *Gainett, G., Sharma, P.P., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2014) Walk it off: Predictive power of appendicular characters toward inference of higher-level relationships in Laniatores (Arachnida: Opiliones). Cladistics 30:120-138. [PDF] (*undergraduate co-author)