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Selected publications
(at UW-Madison)
(at UW-Madison)
Underlined authors are Sharma group members or affiliates.
For complete bibliography, please download CVs on the personnel pages.
Peer reviewed
119. *Gainett, G., *Klementz, B.C., Setton, E.V.W., Simian, C., Iuri, H.A., Edgecombe, G.D., Peretti, A.V., Sharma, P.P. (in review) A plurality of morphological characters need not equate with phylogenetic accuracy: A rare genomic change refutes the placement of Solifugae and Pseudoscorpiones in Haplocnemata. (*equal author contribution)
118. Setton, EV.W., Ballesteros, J.A., Blaszczyk, P.O., Klementz, B.C., Sharma, P.P. (in review) A taxon-restricted duplicate of Iroquois3 is required for patterning the spider waist. [Biorxiv preprint: https://doi.org/10.1101/2023.08.08.552527] [Preprint PDF]
117. Warburg, S., Aharon, S., Armiach Steinpress, I., Sharma, P.P., Harms, D., Gavish-Regev, E. (in review) Pseudoscorpions of Israel: Annotated checklist and key, with new records of two families (Arachnida: Pseudoscorpiones).
116. Klementz, B.C., Sharma, P.P. (in review) New species of Paktongius Suzuki, 1969 (Opiliones: Laniatores: Assamiidae) and convergent evolution of the gonyleptid habitus in Southeast Asian Assamiidae.
115. de Miranda, G.S., Kulkarni, S.S., Tagliatela, J., Baker, C.M., Giupponi, A.P.L., Labarque, F.M., Gavish-Regev, E., Rix, M.G., Carvalho, L.S., Fusari, L.M., Wood, H.M., Sharma, P.P. (in revision) The rediscovery of a relict unlocks the first global phylogeny of whip spiders (Amblypygi). [Biorxiv preprint: https://doi.org/10.1101/2022.04.26.489547] [Preprint PDF]
114. Kulkarni, S.S., *Steiner, H.G., Garcia, E.L., Iuri, H., Jones, R.R., Ballesteros, J.A., Gainett, G., Graham, M.R., Harms, D., Lyle, R., Ojanguren-Affilastro, A.A., Santibañez-López, C.E., Silva de Miranda, G., Cushing, P.E., Gavish-Regev, E., Sharma, P.P. (2023) Neglected no longer: Phylogenomic resolution of higher- level relationships in Solifugae. iScience 26:107684. (*undergraduate co-author) [PDF]
113. Santibáñez-López, C.E., Ojanguren-Affilastro, A.A., Graham, M.R., Sharma, P.P. (2023) Congruence between ultraconserved element-based matrices and phylotranscriptomic datasets in the scorpion tree of life. Cladistics (doi: 10.1111/cla.12551). [PDF]
112. Sharma, P.P. (2023) Impacts of whole genome duplication on the evolution of the arachnids. Integrative and Comparative Biology icad050. [PDF]
111. Sharma, P.P. (2023) Duplication and evolution of Hox clusters in Chelicerata (Arthropoda). In: Ferrier, D.E.K., (ed.). Hox modules in Evolution and Development. Boca Raton, FL: CRC Press, Taylor Francis Group, pp. 77-102. [PDF]
110. Gainett, G., *Klementz, B.C., *Błaszczyk, P.O., Bruce, H.S., Patel, N.D., Sharma, P.P. (2023) Dual functions of labial resolve the Hox logic of chelicerate head segments. Molecular Biology and Evolution 40:msad037. (*undergraduate co-author) [PDF]
109. Aharon, S., Ballesteros, J.A., Gainett, G., Sharma, P.P., Hawlena, D., Gavish-Regev, E. (2023) In the land of the blind: Exceptional subterranean speciation of cryptic troglobitic spiders of the genus Tegenaria (Araneae: Agelenidae) in Israel. Molecular Phylogenetics and Evolution 182:107705.
108. Palmieri, L., Giribet, G., Sharma, P.P. (2023) Too early for the ferry: The biogeographic history of the Assamiidae of southeast Asia (Chelicerata: Opiliones, Laniatores). Molecular Phylogenetics and Evolution 78:107647. [PDF]
107. Palmieri, L., Pavarini, R., Sharma, P.P. (2022) Draft genome sequence of Candidatus Nardonella dryophthoridicola NARMHE1, endosymbiont of Metamasius hemipterus (Coleoptera, Curculionidae, Dryophthorinae). Microbiology Resource Announcements 11:11. [PDF]
106. Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Tarasov, P.E., Ren, D. (2022) Four new Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber. Palaeoworld (doi:10.1016/j.palwor.2022.06.006) [PDF]
105. Palmieri Rocha, L., Chamorro, M.L., Sharma, P.P. (2022) Phylogenetic assessment of the Metamasius hemipterus species complex (Coleoptera, Curculionidae, Dryophthorinae). Molecular Phylogenetics and Evolution 175:107589. [PDF]
104. Baker, C.M., Ballesteros, J.A., Aharon, S., Gainett, G., Armiach Steinpress, I., Wizen, G., Sharma, P.P., Gavish-Regev, E. (2022) Recent speciation and phenotypic plasticity within a parthenogenetic lineage of levantine whip spiders (Chelicerata: Amblypygi: Charinidae). Molecular Phylogenetics and Evolution 175:107560. [PDF]
103. Santibáñez-López, C.E., Aharon, S., Ballesteros, J.A., Gainett, G., Baker, C.M., González-Santillán, E., Harvey, M.S., Hassan, M.K., Abu-Almaaty, A.H., Aldeyarbi, S.M., Monod, L., Ojanguren-Affilastro, A., Raven, R.J., Pinto-da-Roca, R., Zvik, Y., Gavish-Revev, E., Sharma, P.P. (2022) Phylogenomics of scorpions reveal contemporaneous diversification of scorpion mammalian predators and mammal-specific sodium channel toxins. Systematic Biology. (doi: 10.1093/sysbio/syac021) [PDF]
102. *Gainett, G., *^Crawford, A.R., ^Klementz, B.C., ^So, C., Baker, C.M., Setton, E.V.W., Sharma, P.P. (2022) Eggs to longlegs: Embryonic staging of the harvestman Phalangium opilio (Opiliones), an emerging model arachnid. Frontiers in Zoology 19:11. [PDF] (*equal author contribution) (^undergraduate co-author)
101. Ballesteros, J.A., Santibáñez-López, C.E., Baker, C.M., Benavides, L.R., Cunha, T.J., Gainett, G., Ontano, A.Z., Setton, E.V.W., Arango, C.P., Gavish-Regev, E., Harvey, M.S., Wheeler, W.C., Hormiga, G., Giribet, G., Sharma, P.P. (2022) Comprehensive species sampling and sophisticated algorithmic approaches refute the monophyly of Arachnida. Molecular Biology and Evolution 39(2):msac021 [PDF] [Featured in the New York Times]
100. Ontano, A.Z., *Steiner, H.G., Sharma P.P. (2022) How many long branch orders occur in Chelicerata? Opposing effects of Palpigradi and Opilioacariformes on phylogenetic stability. Molecular Phylogenetics and Evolution 168:107378. [PDF] (*undergraduate co-author)
99. Sharma, P.P., Ballesteros, J.A., Santibáñez-López, C.E. (2021) What is an “arachnid”? Consensus, consilience, and confirmation bias in the phylogenetics of Chelicerata. Diversity 13:568. [PDF] [Highlighted by the journal]
98. Gainett, G., González, V.L., Ballesteros, J.A., Setton, E.V.W., Baker, C.M., Barolo Gargiulo, L., Santibáñez-López, C.E., Coddington, J.A., Sharma, P.P. (2021) The genome of a daddy-long-legs (Opiliones) illuminates the evolution of arachnid appendages. Proceedings of the Royal Society B: Biological Sciences 288:20211168. [PDF] [Featured on the journal cover] [Featured in Nature] [Featured in Science] [Featured in the New York Times] [Featured in The Atlantic] [Featured on National Public Radio]
97. Cotoras, D.D., Castanheira, P.d.S., Sharma, P.P.. (2021) A cheliceral axial duplication in the long-jawed orb-weaving spider Tetragnatha versicolor (Araneae: Tetragnathidae). Development Genes and Evolution (doi: 10.1007/s00427-021-00678-9) [PDF]
96. Baudouin-González, L., Schoenaeur, A., Harper, A., Blakely, G., Seiter, M., Arif, S., Summer-Rooney, L., Russel, S., Sharma, P.P., McGregor, A.P. (2021) The evolution of Sox gene repertoires and regulation of segmentation in arachnids. Molecular Biology and Evolution 38:3153–3169 [PDF]
95. Setton, E.V.W., Sharma, P.P. (2021) A conserved role for arrow in posterior axis patterning across Arthropoda. Developmental Biology 475:91-105. [PDF]
94. Ontano, A.Z., Gainett, G., Aharon, S., Ballesteros, J.A., Benavides, L.R., *Corbett, K.F., Gavish-Regev, E., Harvey, M.S., Monsma, S., Santibáñez-López, C.E., Setton, E.V.W., *Zehms, J.T., Zeh, J.A., Zeh, D.W., Sharma, P.P. (2021) Taxonomic sampling and rare genomic changes overcome long-branch attraction in the phylogenetic placement of pseudoscorpions. Molecular Biology and Evolution 38:2446-2467. (*undergraduate co-author) [PDF]
93. *Ballesteros, J.A., *Setton, E.V.W., *Santibáñez-López, C.E., Arango, C.P., Brenneis, G., Brix, S., Cano-Sánchez, E., ^Corbett, K.F., ^Dandouch, M., Dilly, G.F., Eleaume, M., Gainett, G., Gallut, C., ^McAtee, S., ^McIntyre, L., Moran, A.L., ^Moran, R., López-González, P.J., Scholtz, G., ^Williamson, C., Woods, H.A., ^Zehms, J.T., Wheeler, W.C., Sharma, P.P. (2020) Phylogenomic resolution of sea spider diversification through integration of multiple data classes. Molecular Biology and Evolution 38(2):686-701. [PDF] (*Equal author contribution) (^undergraduate co-author) [Featured in Science]
92. Giribet, G., Baker, C.M., Sharma, P.P. (2021) A revised phylogeny of the New Caledonian endemic genus Troglosiro (Opiliones : Cyphophthalmi : Troglosironidae) with the description of four new species. Invertebrate Systematics 35:59-89. [PDF]
91. Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Ren, D., Shi C. (2021) Laniatorid harvestmen (Arachnida) from Late Cretaceous Burmese amber. Cretaceous Research 119:104703. [PDF]
90. Dehnert, G.K., Freitas, M., Sharma, P.P., Barry, T., Karasov, W.H. (2021) Impacts of sub-chronic exposure to a commercial 2,4-D herbicide on developmental stages of multiple freshwater fish species. Chemosphere 263:127638. [PDF]
89. *Gainett, G., *Ballesteros, J.A., Kanzler, C.R., ^Zehms, J.T., ^Zern, J.M., Aharon, S., Gavish-Regev, E., Sharma, P.P. (2020) Systemic paralogy and function of retinal determination network homologs in arachnids. (*Equal author contribution) BMC Genomics 21:811. [PDF] (^undergraduate co-author)
88. Gainett, G., Willemart, R.H., Giribet, G., Sharma, P.P. (2020) Convergent evolution of sexually dimorphic glands in an amphi-Pacific harvestman family. Invertebrate Systematics 34:871-892. [PDF]
87. Brenneis, G., Arango, C.P., Sharma, P.P., Schwentner, M. (2020) The more the merrier: Unparalleled sympatric species richness in a sea spider genus (Pycnogonida: Callipallenidae: Pallenella) from Tasmanian waters. Invertebrate Systematics 34:837-870. [PDF] [Featured on the journal cover]
86. Gainett, G., Sharma, P.P. (2020) New genomic resources and toolkits for developmental study of whip spiders (Amblypygi) provide insights into arachnid genome evolution and antenniform leg patterning. EvoDevo 11:18. [PDF]
85. *Schmidt, S.M., Clouse, R.M., Sharma, P.P. (2020) A new Miopsalis from Mindanao supports a biogeographic umbilicus between Borneo and the southern Philippines (Arachnida: Opiliones: Cyphophthalmi: Stylocellidae). Zootaxa 4779:379-390. [PDF] (*undergraduate co-author)
84. Cano-Sánchez, E., Sharma, P.P., López-González, P.J. (2020) Postembryonic development of Nymphon australe Hodgson, 1902 (Pycnogonida, Nymphonidae) from Antarctica. Polar Biology 43:207-223. [PDF]
83. Santibanez-López, C.E., Ojanguren-Affilastro, A.A., Sharma, P.P. (2020) Another one bites the dust: taxonomic sampling of a key genus in phylogenomic datasets reveals more non-monophyletic groups in traditional scorpion classification. Invertebrate Systematics 34:133-143. [PDF]
82. Nolan, E.D., Santibáñez-López, C.E., Sharma, P.P. (2020) Developmental gene expression as a phylogenetic data class: Support for the monophyly of Arachnopulmonata. Development Genes and Evolution 230:137-153. [PDF] [Featured on the journal cover]
81.*Ballesteros, J.A., *Santibáñez-López, C.E, Kováč, L', Gavish-Regev, E., Sharma, P.P. (2019) Ordered phylogenomic subsampling enables diagnosis of systematic errors in the placement of the enigmatic arachnid order Palpigradi. Proceedings of the Royal Society of London B: Biological Sciences 286:20192426. [PDF] (*Equal author contribution)
80. Santibáñez-López, C.E, Graham, M.R., Sharma, P.P., Ortiz, E., Possani, L. (2019) Comparative transcriptomic analysis reveals evolutionary conservation and selective pressures in hadrurid scorpion toxins. Toxins 11:637. [PDF] [Featured on the journal cover]
79. Aharon, S., Ballesteros, J.A., *Crawford, A.C., *Friske, K., Guilherme, G., Langford, B., Santibáñez-López, C.E., Ya'aran, S., Gavish-Regev, E. & Sharma, P.P. (2019) The anatomy of an unstable node: A Levantine relict precipitates phylogenomic dissolution of higher-level relationships of the armored harvestmen (Arachnida: Opiliones: Laniatores). Invertebrate Systematics 33:697-717. [PDF] (*undergraduate co-author) [Featured on the journal cover]
78. Santibáñez-López, C.E., González-Santillán, E., Monod, L., Sharma, P.P. (2019) Phylogenomics facilitates stable scorpion systematics: Reassessing the relationships of Vaejovidae and a new high-level classification of Scorpiones (Arachnida). Molecular Phylogenetics and Evolution 135:22-30. [PDF]
77. Setton, E.V.W., Hendrixson, B.E. & Sharma, P.P. (2019) Embryogenesis in a Colorado population of Aphonopelma hentzi (Girard, 1852), a promising system for the study of development (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology 47:209-216. [PDF] [Featured on the journal cover]
76. Gainett, G., Sharma, P.P., Fernandes, N., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2019) Evolution of a sensory cluster on the legs of Opiliones (Arachnida) informs multi-level phylogenetic relationships. Zoological Journal of the Linnean Society 187:143-165. [PDF]
75. Ballesteros, J.A. & Sharma, P.P. (2019) A critical appraisal of the placement of Xiphosura (Chelicerata) with account of known sources of phylogenetic error. Systematic Biology 68:896-917. [PDF] [Featured on the journal cover] [Featured in National Geographic]
74. Sharma, P.P. (2019) Integrating morphology and phylogenomics supports a terrestrial origin of insect flight. Proceedings of the National Academy of Sciences of the USA 116:2796-2798. [PDF]
73. *Schmidt, S.M., Buenavente, P.A.C., Blatchley, D.D., Diesmos, A.C., Diesmos, M.L., General, D.E.M., Mohagan, A.B., Mohagan, D.J., Clouse, R.M. & Sharma, P.P. (2019) A new species of Tithaeidae (Arachnida: Opiliones: Laniatores) from Mindanao reveals contemporaneous colonization of the Philippines by Sunda Shelf opiliofauna. Invertebrate Systematics 33:237-251. [PDF]
72. *March, L.E., *Smaby, R.M., *Setton, E.V.W. & Sharma, P.P. (2018) The evolution of selector gene function: Expression dynamics and regulatory interactions of tiptop/teashirt across Arthropoda. Evolution & Development 20:219-232. [PDF] (*undergraduate co-author)
71. Santibañez López, C.E., Kriebel, R., Ballesteros, J.A., Rush, N., Witter, Z., Williams, J., Janies, D.A. & Sharma, P.P. (2018) Integration of phylogenomics and molecular modeling reveals lineage-specific diversification of toxins in scorpions. PeerJ 5902:1-13. [PDF]
70. Zhang, C., Zhang F. & Sharma, P.P. (2018) Two new species of Petrobunus Sharma & Giribet, 2011 from China (Opiliones: Laniatores: Petrobunidae). Zootaxa 4524:51-64. [PDF]
69. Leite, D.J., Baudouin-Gonzalez, L., Iwasaki-Yokozawa, S., Lozano-Fernandez, J., Turetzek, N., Akiyama-Oda, Y., Pisani, D., Oda, H., Sharma, P.P. & McGregor, A.P. (2018) Homeobox gene duplication and divergence in arachnids. Molecular Biology and Evolution 35:2240-2253. [PDF]
68. Sharma, P.P. (2018) Chelicerates. Current Biology 28:R774-778. [PDF]
67. Oberski, J.T., Sharma, P.P., Jay, K.R., Coblens, M., Lemon, K.A., Johnson, J.E. & Boyer, S.L. (2018) A dated molecular phylogeny of mite harvestmen (Arachnida: Opiliones: Cyphophthalmi) elucidates ancient diversification dynamics in the Australian Wet Tropics. Molecular Phylogenetics and Evolution 127:813-822. [PDF].
66. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) The sensory equipment of a sandokanid: an extreme case of tarsal reduction in a harvestman (Arachnida, Opiliones). Journal of Morphology (doi: 10.1002/jmor.20843). [PDF]
65. Di, Z., Edgecombe, G.D., Sharma, P.P. (2018) Homeosis in a scorpion supports a telopodal origin of pectines and book lungs. BMC Evolutionary Biology 18:73. [PDF]
64. Santibáñez-López, C.E., Ontano, A.Z., Harvey, M.S., & Sharma, P.P. (2018) Transcriptomic analysis of pseudoscorpion venom reveals a unique cocktail dominated by enzymes and protease inhibitors. Toxins 10:207. [PDF]
63. *Setton, E.V.W., Sharma, P.P. (2018) Cooption of an appendage patterning gene cassette in the head segmentation of arachnids. Proceedings of the National Academy of Sciences of the USA 115:E3491-E3500. [PDF] (*undergraduate co-author)
62. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) Putative adhesive setae on the walking legs of the Paleotropical harvestman Metibalonius sp. (Arachnida: Opiliones: Podoctidae). Journal of Arachnology 46:62-68. [PDF]
61. Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. & Giribet, G. (2018) A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122:37-45. [PDF]
60. Garb, J.E., Sharma, P.P. & Ayoub, N.A. (2018) Recent progress and prospects for advancing arachnid genomics. Current Opinions in Insect Science 25:51-57. [PDF]
59. Lewis, S.H., Quarles, K.A., Yang, Y., Tanguy, M., Frézal, L., Smith, S., Sharma, P.P, Cordaux, R., Gilbert, C., Giraud, I., Collins, D.H., Zamore, P.D., Miska, E.A., Sarkies, P. & Jiggins, F.M. (2018) Pan-arthropod analysis reveals somatic piRNAs as an ancestral defence against transposable elements. Nature Ecology and Evolution 2:174-181. [PDF]
58. Daniels, S.R., Dreyer, M. & Sharma, P.P. (2017) Contrasting the population genetic structure of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsis) in forest fragments along the south eastern Cape, South Africa. Invertebrate Systematics 31:781-796. [PDF]
57. Sharma, P.P. (2017) Chelicerates and the conquest of land: A view of arachnid origins through an evodevo spyglass. Integrative and Comparative Biology 57, 510-522. [PDF]
56. *Schwager, E.E., *Sharma, P.P., *Clarke, T., *Leite, D.J., Wierschin, T., Pechmann M., Akiyama-Oda, Y., Esposito, L., Arensburger, P., Bechsgaard, J., Bilde, T., Buffry, A., Chao, H., Dinh, H., Doddapaneni, H., Dugan, S., Eibner C., Extavour, C.G., Funch, P., Garb, J., Gonzalez, V.L., Griffiths-Jones, S., Han, Y., Hayashi, C., Hilbrant, M., Hughes, D.S.T., Janssen, R., Lee, S.L., Maeso, I., Murali, S.C., Muzny, D.M., da Fonseca, R.N., Qu, J., Ronshaugen, M., Schomburg, C., Schönauer, A., Stollewerk, A., Torres-Oliva, M., Turetzek, N., Vanthournout, B., Werren, J., Wolff, C., Worley, K.C., Gibbs, R.A., Coddington, J., Oda, H., Stanke, M., Ayoub, N.A., Damen, W.G.M., Prpic, N.-M., Flot, J.-F., Posnien, N., Richards, S., & McGregor, A.P. (2017) The house spider genome reveals a whole genome duplication during arachnid evolution. BMC Biology 15:62. [PDF] (*equal author contribution)
55. *Setton, E.V.W., *March, L.E., *Nolan, E.D., Jones, T.E., Cho, H., Wheeler, W.C., Extavour, C.G. & Sharma, P.P. (2017) Expression and function of spineless orthologs correlate with distal deutocerebral appendage morphology in Arthropoda. Developmental Biology 430:224-236. (*undergraduate co-author) [PDF]
54. Clouse, R.M., Branstetter, M.G., Buenavente, P.A.C., Crowley, L.M., Czekanski-Moir, J., General, D.E.M., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Sharma, P.P. & Wheeler, W.C. (2017) First molecular phylogeny of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44:2660-2672. [PDF] [Featured on the journal cover]
53. Sharma P.P., Oberski, J.T., Santiago, M.A., Ricardo Kriebel, *Savana M. Lipps, Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). Molecular Phylogenetics and Evolution. (*undergraduate co-author) (doi: 10.1016/j.ympev.2017.03.026) [PDF]
52. *Santibañez-López, C., *Kriebel, R. & Sharma, P.P. (2017) eadem figura manet: Measuring morphological convergence in diplocentrid scorpions (Arachnida: Scorpiones: Diplocentridae) under a multilocus phylogenetic framework. Invertebrate Systematics. Invertebrate Systematics 31:233-248. [PDF] (*equal author contribution) [Featured on the journal cover]
51. Fernández, R., Sharma P.P., Tourinho, A.L. & Giribet, G. (2017) The Opiliones Tree of Life: shedding light on harvestmen relationships through transcriptomics. Proceedings of the Royal Society of London B: Biological Sciences 284:20162340. [PDF]
50. Sharma P.P., *Santiago, M.A., Kriebel, R., *Lipps, S.M., Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) reveals the disutility of subfamily nomenclature and traditional character systems in armored harvestman systematics. Molecular Phylogenetics and Evolution 106:164-173. [PDF] (*undergraduate co-author)
49. *Jay, K.R., Popkin-Hall, Z.R., *Coblens, M.J., *Oberski, J.T., Sharma, P.P. & Boyer, S.L. (2016) New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot. Zookeys 586:37-93. [PDF] (*undergraduate co-author)
48. Iyer, H., Issigonis, M., Sharma, P.P., Extavour, C.G. & Newmark, P.A. (2016) A novel pre-meiotic function for boule in the planarian Schmidtea mediterranea. Proceedings of the National Academy of Sciences of the USA 113:E3509-E3518. (doi: 10.1073/pnas.1521341113) [PDF]
47. Sharma, P.P., Clouse, R.M. & Wheeler, W.C. (2016) Hennig's semaphoront concept and the use of ontogenetic stages in phylogenetic reconstruction. Cladistics. (doi: 10.1111/cla.12156) [PDF]
46. Daniels, S.R., Kunaka, C., Klaus, S. & Sharma, P.P. (2016) Unmasking alpha diversity, cladogenesis and biogeographical patterning in an ancient panarthropod lineage (Onychophora: Peripatopsidae: Opisthopatus cinctipes) with the description of five novel species. Cladistics. (doi: 10.1111/cla.12154) [PDF]
45. Giribet, G., Boyer, S.L., Baker, C.M., Fernández, R., Sharma P.P., de Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. & Griswold, C.E. (2016) A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178:523-545. [PDF]
44. Pérez-González, A., Sharma, P.P. & Proud, D.N. (2016) Morphological tricks and blessed genitalia: rectifying the familial placement of Fijicolana tuberculata (Opiliones: Laniatores: Zalmoxidae). Zootaxa 4061:253-260. [PDF]
43. Clouse, R.M., Sharma, P.P., *Stuart, J.C., Davis, L.R., Giribet, G., Sarah, S.L. & Wheeler, W.C. (2016) Phylogeography of the harvestman Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox. Organisms, Diversity and Evolution 16:167-184. [PDF] (*undergraduate co-author)
42. Sharma, P.P., *Santiago, M.A., González-Santillán, E., Monod, L. & Wheeler, W.C. (2015) Evidence of duplicated Hox genes in the most recent common ancestor of extant scorpions. Evolution and Development 17:347-355. [PDF] (*undergraduate co-author)
For complete bibliography, please download CVs on the personnel pages.
Peer reviewed
119. *Gainett, G., *Klementz, B.C., Setton, E.V.W., Simian, C., Iuri, H.A., Edgecombe, G.D., Peretti, A.V., Sharma, P.P. (in review) A plurality of morphological characters need not equate with phylogenetic accuracy: A rare genomic change refutes the placement of Solifugae and Pseudoscorpiones in Haplocnemata. (*equal author contribution)
118. Setton, EV.W., Ballesteros, J.A., Blaszczyk, P.O., Klementz, B.C., Sharma, P.P. (in review) A taxon-restricted duplicate of Iroquois3 is required for patterning the spider waist. [Biorxiv preprint: https://doi.org/10.1101/2023.08.08.552527] [Preprint PDF]
117. Warburg, S., Aharon, S., Armiach Steinpress, I., Sharma, P.P., Harms, D., Gavish-Regev, E. (in review) Pseudoscorpions of Israel: Annotated checklist and key, with new records of two families (Arachnida: Pseudoscorpiones).
116. Klementz, B.C., Sharma, P.P. (in review) New species of Paktongius Suzuki, 1969 (Opiliones: Laniatores: Assamiidae) and convergent evolution of the gonyleptid habitus in Southeast Asian Assamiidae.
115. de Miranda, G.S., Kulkarni, S.S., Tagliatela, J., Baker, C.M., Giupponi, A.P.L., Labarque, F.M., Gavish-Regev, E., Rix, M.G., Carvalho, L.S., Fusari, L.M., Wood, H.M., Sharma, P.P. (in revision) The rediscovery of a relict unlocks the first global phylogeny of whip spiders (Amblypygi). [Biorxiv preprint: https://doi.org/10.1101/2022.04.26.489547] [Preprint PDF]
114. Kulkarni, S.S., *Steiner, H.G., Garcia, E.L., Iuri, H., Jones, R.R., Ballesteros, J.A., Gainett, G., Graham, M.R., Harms, D., Lyle, R., Ojanguren-Affilastro, A.A., Santibañez-López, C.E., Silva de Miranda, G., Cushing, P.E., Gavish-Regev, E., Sharma, P.P. (2023) Neglected no longer: Phylogenomic resolution of higher- level relationships in Solifugae. iScience 26:107684. (*undergraduate co-author) [PDF]
113. Santibáñez-López, C.E., Ojanguren-Affilastro, A.A., Graham, M.R., Sharma, P.P. (2023) Congruence between ultraconserved element-based matrices and phylotranscriptomic datasets in the scorpion tree of life. Cladistics (doi: 10.1111/cla.12551). [PDF]
112. Sharma, P.P. (2023) Impacts of whole genome duplication on the evolution of the arachnids. Integrative and Comparative Biology icad050. [PDF]
111. Sharma, P.P. (2023) Duplication and evolution of Hox clusters in Chelicerata (Arthropoda). In: Ferrier, D.E.K., (ed.). Hox modules in Evolution and Development. Boca Raton, FL: CRC Press, Taylor Francis Group, pp. 77-102. [PDF]
110. Gainett, G., *Klementz, B.C., *Błaszczyk, P.O., Bruce, H.S., Patel, N.D., Sharma, P.P. (2023) Dual functions of labial resolve the Hox logic of chelicerate head segments. Molecular Biology and Evolution 40:msad037. (*undergraduate co-author) [PDF]
109. Aharon, S., Ballesteros, J.A., Gainett, G., Sharma, P.P., Hawlena, D., Gavish-Regev, E. (2023) In the land of the blind: Exceptional subterranean speciation of cryptic troglobitic spiders of the genus Tegenaria (Araneae: Agelenidae) in Israel. Molecular Phylogenetics and Evolution 182:107705.
108. Palmieri, L., Giribet, G., Sharma, P.P. (2023) Too early for the ferry: The biogeographic history of the Assamiidae of southeast Asia (Chelicerata: Opiliones, Laniatores). Molecular Phylogenetics and Evolution 78:107647. [PDF]
107. Palmieri, L., Pavarini, R., Sharma, P.P. (2022) Draft genome sequence of Candidatus Nardonella dryophthoridicola NARMHE1, endosymbiont of Metamasius hemipterus (Coleoptera, Curculionidae, Dryophthorinae). Microbiology Resource Announcements 11:11. [PDF]
106. Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Tarasov, P.E., Ren, D. (2022) Four new Laniatorean harvestmen (Arachnida: Opiliones) from mid-Cretaceous Burmese amber. Palaeoworld (doi:10.1016/j.palwor.2022.06.006) [PDF]
105. Palmieri Rocha, L., Chamorro, M.L., Sharma, P.P. (2022) Phylogenetic assessment of the Metamasius hemipterus species complex (Coleoptera, Curculionidae, Dryophthorinae). Molecular Phylogenetics and Evolution 175:107589. [PDF]
104. Baker, C.M., Ballesteros, J.A., Aharon, S., Gainett, G., Armiach Steinpress, I., Wizen, G., Sharma, P.P., Gavish-Regev, E. (2022) Recent speciation and phenotypic plasticity within a parthenogenetic lineage of levantine whip spiders (Chelicerata: Amblypygi: Charinidae). Molecular Phylogenetics and Evolution 175:107560. [PDF]
103. Santibáñez-López, C.E., Aharon, S., Ballesteros, J.A., Gainett, G., Baker, C.M., González-Santillán, E., Harvey, M.S., Hassan, M.K., Abu-Almaaty, A.H., Aldeyarbi, S.M., Monod, L., Ojanguren-Affilastro, A., Raven, R.J., Pinto-da-Roca, R., Zvik, Y., Gavish-Revev, E., Sharma, P.P. (2022) Phylogenomics of scorpions reveal contemporaneous diversification of scorpion mammalian predators and mammal-specific sodium channel toxins. Systematic Biology. (doi: 10.1093/sysbio/syac021) [PDF]
102. *Gainett, G., *^Crawford, A.R., ^Klementz, B.C., ^So, C., Baker, C.M., Setton, E.V.W., Sharma, P.P. (2022) Eggs to longlegs: Embryonic staging of the harvestman Phalangium opilio (Opiliones), an emerging model arachnid. Frontiers in Zoology 19:11. [PDF] (*equal author contribution) (^undergraduate co-author)
101. Ballesteros, J.A., Santibáñez-López, C.E., Baker, C.M., Benavides, L.R., Cunha, T.J., Gainett, G., Ontano, A.Z., Setton, E.V.W., Arango, C.P., Gavish-Regev, E., Harvey, M.S., Wheeler, W.C., Hormiga, G., Giribet, G., Sharma, P.P. (2022) Comprehensive species sampling and sophisticated algorithmic approaches refute the monophyly of Arachnida. Molecular Biology and Evolution 39(2):msac021 [PDF] [Featured in the New York Times]
100. Ontano, A.Z., *Steiner, H.G., Sharma P.P. (2022) How many long branch orders occur in Chelicerata? Opposing effects of Palpigradi and Opilioacariformes on phylogenetic stability. Molecular Phylogenetics and Evolution 168:107378. [PDF] (*undergraduate co-author)
99. Sharma, P.P., Ballesteros, J.A., Santibáñez-López, C.E. (2021) What is an “arachnid”? Consensus, consilience, and confirmation bias in the phylogenetics of Chelicerata. Diversity 13:568. [PDF] [Highlighted by the journal]
98. Gainett, G., González, V.L., Ballesteros, J.A., Setton, E.V.W., Baker, C.M., Barolo Gargiulo, L., Santibáñez-López, C.E., Coddington, J.A., Sharma, P.P. (2021) The genome of a daddy-long-legs (Opiliones) illuminates the evolution of arachnid appendages. Proceedings of the Royal Society B: Biological Sciences 288:20211168. [PDF] [Featured on the journal cover] [Featured in Nature] [Featured in Science] [Featured in the New York Times] [Featured in The Atlantic] [Featured on National Public Radio]
97. Cotoras, D.D., Castanheira, P.d.S., Sharma, P.P.. (2021) A cheliceral axial duplication in the long-jawed orb-weaving spider Tetragnatha versicolor (Araneae: Tetragnathidae). Development Genes and Evolution (doi: 10.1007/s00427-021-00678-9) [PDF]
96. Baudouin-González, L., Schoenaeur, A., Harper, A., Blakely, G., Seiter, M., Arif, S., Summer-Rooney, L., Russel, S., Sharma, P.P., McGregor, A.P. (2021) The evolution of Sox gene repertoires and regulation of segmentation in arachnids. Molecular Biology and Evolution 38:3153–3169 [PDF]
95. Setton, E.V.W., Sharma, P.P. (2021) A conserved role for arrow in posterior axis patterning across Arthropoda. Developmental Biology 475:91-105. [PDF]
94. Ontano, A.Z., Gainett, G., Aharon, S., Ballesteros, J.A., Benavides, L.R., *Corbett, K.F., Gavish-Regev, E., Harvey, M.S., Monsma, S., Santibáñez-López, C.E., Setton, E.V.W., *Zehms, J.T., Zeh, J.A., Zeh, D.W., Sharma, P.P. (2021) Taxonomic sampling and rare genomic changes overcome long-branch attraction in the phylogenetic placement of pseudoscorpions. Molecular Biology and Evolution 38:2446-2467. (*undergraduate co-author) [PDF]
93. *Ballesteros, J.A., *Setton, E.V.W., *Santibáñez-López, C.E., Arango, C.P., Brenneis, G., Brix, S., Cano-Sánchez, E., ^Corbett, K.F., ^Dandouch, M., Dilly, G.F., Eleaume, M., Gainett, G., Gallut, C., ^McAtee, S., ^McIntyre, L., Moran, A.L., ^Moran, R., López-González, P.J., Scholtz, G., ^Williamson, C., Woods, H.A., ^Zehms, J.T., Wheeler, W.C., Sharma, P.P. (2020) Phylogenomic resolution of sea spider diversification through integration of multiple data classes. Molecular Biology and Evolution 38(2):686-701. [PDF] (*Equal author contribution) (^undergraduate co-author) [Featured in Science]
92. Giribet, G., Baker, C.M., Sharma, P.P. (2021) A revised phylogeny of the New Caledonian endemic genus Troglosiro (Opiliones : Cyphophthalmi : Troglosironidae) with the description of four new species. Invertebrate Systematics 35:59-89. [PDF]
91. Bartel, C., Dunlop, J.A., Sharma, P.P., Selden, P.A., Ren, D., Shi C. (2021) Laniatorid harvestmen (Arachnida) from Late Cretaceous Burmese amber. Cretaceous Research 119:104703. [PDF]
90. Dehnert, G.K., Freitas, M., Sharma, P.P., Barry, T., Karasov, W.H. (2021) Impacts of sub-chronic exposure to a commercial 2,4-D herbicide on developmental stages of multiple freshwater fish species. Chemosphere 263:127638. [PDF]
89. *Gainett, G., *Ballesteros, J.A., Kanzler, C.R., ^Zehms, J.T., ^Zern, J.M., Aharon, S., Gavish-Regev, E., Sharma, P.P. (2020) Systemic paralogy and function of retinal determination network homologs in arachnids. (*Equal author contribution) BMC Genomics 21:811. [PDF] (^undergraduate co-author)
88. Gainett, G., Willemart, R.H., Giribet, G., Sharma, P.P. (2020) Convergent evolution of sexually dimorphic glands in an amphi-Pacific harvestman family. Invertebrate Systematics 34:871-892. [PDF]
87. Brenneis, G., Arango, C.P., Sharma, P.P., Schwentner, M. (2020) The more the merrier: Unparalleled sympatric species richness in a sea spider genus (Pycnogonida: Callipallenidae: Pallenella) from Tasmanian waters. Invertebrate Systematics 34:837-870. [PDF] [Featured on the journal cover]
86. Gainett, G., Sharma, P.P. (2020) New genomic resources and toolkits for developmental study of whip spiders (Amblypygi) provide insights into arachnid genome evolution and antenniform leg patterning. EvoDevo 11:18. [PDF]
85. *Schmidt, S.M., Clouse, R.M., Sharma, P.P. (2020) A new Miopsalis from Mindanao supports a biogeographic umbilicus between Borneo and the southern Philippines (Arachnida: Opiliones: Cyphophthalmi: Stylocellidae). Zootaxa 4779:379-390. [PDF] (*undergraduate co-author)
84. Cano-Sánchez, E., Sharma, P.P., López-González, P.J. (2020) Postembryonic development of Nymphon australe Hodgson, 1902 (Pycnogonida, Nymphonidae) from Antarctica. Polar Biology 43:207-223. [PDF]
83. Santibanez-López, C.E., Ojanguren-Affilastro, A.A., Sharma, P.P. (2020) Another one bites the dust: taxonomic sampling of a key genus in phylogenomic datasets reveals more non-monophyletic groups in traditional scorpion classification. Invertebrate Systematics 34:133-143. [PDF]
82. Nolan, E.D., Santibáñez-López, C.E., Sharma, P.P. (2020) Developmental gene expression as a phylogenetic data class: Support for the monophyly of Arachnopulmonata. Development Genes and Evolution 230:137-153. [PDF] [Featured on the journal cover]
81.*Ballesteros, J.A., *Santibáñez-López, C.E, Kováč, L', Gavish-Regev, E., Sharma, P.P. (2019) Ordered phylogenomic subsampling enables diagnosis of systematic errors in the placement of the enigmatic arachnid order Palpigradi. Proceedings of the Royal Society of London B: Biological Sciences 286:20192426. [PDF] (*Equal author contribution)
80. Santibáñez-López, C.E, Graham, M.R., Sharma, P.P., Ortiz, E., Possani, L. (2019) Comparative transcriptomic analysis reveals evolutionary conservation and selective pressures in hadrurid scorpion toxins. Toxins 11:637. [PDF] [Featured on the journal cover]
79. Aharon, S., Ballesteros, J.A., *Crawford, A.C., *Friske, K., Guilherme, G., Langford, B., Santibáñez-López, C.E., Ya'aran, S., Gavish-Regev, E. & Sharma, P.P. (2019) The anatomy of an unstable node: A Levantine relict precipitates phylogenomic dissolution of higher-level relationships of the armored harvestmen (Arachnida: Opiliones: Laniatores). Invertebrate Systematics 33:697-717. [PDF] (*undergraduate co-author) [Featured on the journal cover]
78. Santibáñez-López, C.E., González-Santillán, E., Monod, L., Sharma, P.P. (2019) Phylogenomics facilitates stable scorpion systematics: Reassessing the relationships of Vaejovidae and a new high-level classification of Scorpiones (Arachnida). Molecular Phylogenetics and Evolution 135:22-30. [PDF]
77. Setton, E.V.W., Hendrixson, B.E. & Sharma, P.P. (2019) Embryogenesis in a Colorado population of Aphonopelma hentzi (Girard, 1852), a promising system for the study of development (Araneae, Mygalomorphae, Theraphosidae). Journal of Arachnology 47:209-216. [PDF] [Featured on the journal cover]
76. Gainett, G., Sharma, P.P., Fernandes, N., Pinto-da-Rocha, R., Giribet, G. & Willemart, R.H. (2019) Evolution of a sensory cluster on the legs of Opiliones (Arachnida) informs multi-level phylogenetic relationships. Zoological Journal of the Linnean Society 187:143-165. [PDF]
75. Ballesteros, J.A. & Sharma, P.P. (2019) A critical appraisal of the placement of Xiphosura (Chelicerata) with account of known sources of phylogenetic error. Systematic Biology 68:896-917. [PDF] [Featured on the journal cover] [Featured in National Geographic]
74. Sharma, P.P. (2019) Integrating morphology and phylogenomics supports a terrestrial origin of insect flight. Proceedings of the National Academy of Sciences of the USA 116:2796-2798. [PDF]
73. *Schmidt, S.M., Buenavente, P.A.C., Blatchley, D.D., Diesmos, A.C., Diesmos, M.L., General, D.E.M., Mohagan, A.B., Mohagan, D.J., Clouse, R.M. & Sharma, P.P. (2019) A new species of Tithaeidae (Arachnida: Opiliones: Laniatores) from Mindanao reveals contemporaneous colonization of the Philippines by Sunda Shelf opiliofauna. Invertebrate Systematics 33:237-251. [PDF]
72. *March, L.E., *Smaby, R.M., *Setton, E.V.W. & Sharma, P.P. (2018) The evolution of selector gene function: Expression dynamics and regulatory interactions of tiptop/teashirt across Arthropoda. Evolution & Development 20:219-232. [PDF] (*undergraduate co-author)
71. Santibañez López, C.E., Kriebel, R., Ballesteros, J.A., Rush, N., Witter, Z., Williams, J., Janies, D.A. & Sharma, P.P. (2018) Integration of phylogenomics and molecular modeling reveals lineage-specific diversification of toxins in scorpions. PeerJ 5902:1-13. [PDF]
70. Zhang, C., Zhang F. & Sharma, P.P. (2018) Two new species of Petrobunus Sharma & Giribet, 2011 from China (Opiliones: Laniatores: Petrobunidae). Zootaxa 4524:51-64. [PDF]
69. Leite, D.J., Baudouin-Gonzalez, L., Iwasaki-Yokozawa, S., Lozano-Fernandez, J., Turetzek, N., Akiyama-Oda, Y., Pisani, D., Oda, H., Sharma, P.P. & McGregor, A.P. (2018) Homeobox gene duplication and divergence in arachnids. Molecular Biology and Evolution 35:2240-2253. [PDF]
68. Sharma, P.P. (2018) Chelicerates. Current Biology 28:R774-778. [PDF]
67. Oberski, J.T., Sharma, P.P., Jay, K.R., Coblens, M., Lemon, K.A., Johnson, J.E. & Boyer, S.L. (2018) A dated molecular phylogeny of mite harvestmen (Arachnida: Opiliones: Cyphophthalmi) elucidates ancient diversification dynamics in the Australian Wet Tropics. Molecular Phylogenetics and Evolution 127:813-822. [PDF].
66. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) The sensory equipment of a sandokanid: an extreme case of tarsal reduction in a harvestman (Arachnida, Opiliones). Journal of Morphology (doi: 10.1002/jmor.20843). [PDF]
65. Di, Z., Edgecombe, G.D., Sharma, P.P. (2018) Homeosis in a scorpion supports a telopodal origin of pectines and book lungs. BMC Evolutionary Biology 18:73. [PDF]
64. Santibáñez-López, C.E., Ontano, A.Z., Harvey, M.S., & Sharma, P.P. (2018) Transcriptomic analysis of pseudoscorpion venom reveals a unique cocktail dominated by enzymes and protease inhibitors. Toxins 10:207. [PDF]
63. *Setton, E.V.W., Sharma, P.P. (2018) Cooption of an appendage patterning gene cassette in the head segmentation of arachnids. Proceedings of the National Academy of Sciences of the USA 115:E3491-E3500. [PDF] (*undergraduate co-author)
62. Gainett, G., Sharma, P.P., Giribet, G., Willemart, R.H. (2018) Putative adhesive setae on the walking legs of the Paleotropical harvestman Metibalonius sp. (Arachnida: Opiliones: Podoctidae). Journal of Arachnology 46:62-68. [PDF]
61. Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. & Giribet, G. (2018) A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122:37-45. [PDF]
60. Garb, J.E., Sharma, P.P. & Ayoub, N.A. (2018) Recent progress and prospects for advancing arachnid genomics. Current Opinions in Insect Science 25:51-57. [PDF]
59. Lewis, S.H., Quarles, K.A., Yang, Y., Tanguy, M., Frézal, L., Smith, S., Sharma, P.P, Cordaux, R., Gilbert, C., Giraud, I., Collins, D.H., Zamore, P.D., Miska, E.A., Sarkies, P. & Jiggins, F.M. (2018) Pan-arthropod analysis reveals somatic piRNAs as an ancestral defence against transposable elements. Nature Ecology and Evolution 2:174-181. [PDF]
58. Daniels, S.R., Dreyer, M. & Sharma, P.P. (2017) Contrasting the population genetic structure of two velvet worm taxa (Onychophora: Peripatopsidae: Peripatopsis) in forest fragments along the south eastern Cape, South Africa. Invertebrate Systematics 31:781-796. [PDF]
57. Sharma, P.P. (2017) Chelicerates and the conquest of land: A view of arachnid origins through an evodevo spyglass. Integrative and Comparative Biology 57, 510-522. [PDF]
56. *Schwager, E.E., *Sharma, P.P., *Clarke, T., *Leite, D.J., Wierschin, T., Pechmann M., Akiyama-Oda, Y., Esposito, L., Arensburger, P., Bechsgaard, J., Bilde, T., Buffry, A., Chao, H., Dinh, H., Doddapaneni, H., Dugan, S., Eibner C., Extavour, C.G., Funch, P., Garb, J., Gonzalez, V.L., Griffiths-Jones, S., Han, Y., Hayashi, C., Hilbrant, M., Hughes, D.S.T., Janssen, R., Lee, S.L., Maeso, I., Murali, S.C., Muzny, D.M., da Fonseca, R.N., Qu, J., Ronshaugen, M., Schomburg, C., Schönauer, A., Stollewerk, A., Torres-Oliva, M., Turetzek, N., Vanthournout, B., Werren, J., Wolff, C., Worley, K.C., Gibbs, R.A., Coddington, J., Oda, H., Stanke, M., Ayoub, N.A., Damen, W.G.M., Prpic, N.-M., Flot, J.-F., Posnien, N., Richards, S., & McGregor, A.P. (2017) The house spider genome reveals a whole genome duplication during arachnid evolution. BMC Biology 15:62. [PDF] (*equal author contribution)
55. *Setton, E.V.W., *March, L.E., *Nolan, E.D., Jones, T.E., Cho, H., Wheeler, W.C., Extavour, C.G. & Sharma, P.P. (2017) Expression and function of spineless orthologs correlate with distal deutocerebral appendage morphology in Arthropoda. Developmental Biology 430:224-236. (*undergraduate co-author) [PDF]
54. Clouse, R.M., Branstetter, M.G., Buenavente, P.A.C., Crowley, L.M., Czekanski-Moir, J., General, D.E.M., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Sharma, P.P. & Wheeler, W.C. (2017) First molecular phylogeny of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44:2660-2672. [PDF] [Featured on the journal cover]
53. Sharma P.P., Oberski, J.T., Santiago, M.A., Ricardo Kriebel, *Savana M. Lipps, Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) There is no evidence that Podoctidae carry eggs of their own species: Reply to Machado and Wolff (2017). Molecular Phylogenetics and Evolution. (*undergraduate co-author) (doi: 10.1016/j.ympev.2017.03.026) [PDF]
52. *Santibañez-López, C., *Kriebel, R. & Sharma, P.P. (2017) eadem figura manet: Measuring morphological convergence in diplocentrid scorpions (Arachnida: Scorpiones: Diplocentridae) under a multilocus phylogenetic framework. Invertebrate Systematics. Invertebrate Systematics 31:233-248. [PDF] (*equal author contribution) [Featured on the journal cover]
51. Fernández, R., Sharma P.P., Tourinho, A.L. & Giribet, G. (2017) The Opiliones Tree of Life: shedding light on harvestmen relationships through transcriptomics. Proceedings of the Royal Society of London B: Biological Sciences 284:20162340. [PDF]
50. Sharma P.P., *Santiago, M.A., Kriebel, R., *Lipps, S.M., Buenavente, P.A.C., Diesmos, A.C., Janda, M., Boyer, S.L., Clouse, R.M. & Wheeler, W.C. (2017) A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) reveals the disutility of subfamily nomenclature and traditional character systems in armored harvestman systematics. Molecular Phylogenetics and Evolution 106:164-173. [PDF] (*undergraduate co-author)
49. *Jay, K.R., Popkin-Hall, Z.R., *Coblens, M.J., *Oberski, J.T., Sharma, P.P. & Boyer, S.L. (2016) New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot. Zookeys 586:37-93. [PDF] (*undergraduate co-author)
48. Iyer, H., Issigonis, M., Sharma, P.P., Extavour, C.G. & Newmark, P.A. (2016) A novel pre-meiotic function for boule in the planarian Schmidtea mediterranea. Proceedings of the National Academy of Sciences of the USA 113:E3509-E3518. (doi: 10.1073/pnas.1521341113) [PDF]
47. Sharma, P.P., Clouse, R.M. & Wheeler, W.C. (2016) Hennig's semaphoront concept and the use of ontogenetic stages in phylogenetic reconstruction. Cladistics. (doi: 10.1111/cla.12156) [PDF]
46. Daniels, S.R., Kunaka, C., Klaus, S. & Sharma, P.P. (2016) Unmasking alpha diversity, cladogenesis and biogeographical patterning in an ancient panarthropod lineage (Onychophora: Peripatopsidae: Opisthopatus cinctipes) with the description of five novel species. Cladistics. (doi: 10.1111/cla.12154) [PDF]
45. Giribet, G., Boyer, S.L., Baker, C.M., Fernández, R., Sharma P.P., de Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. & Griswold, C.E. (2016) A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling. Zoological Journal of the Linnean Society 178:523-545. [PDF]
44. Pérez-González, A., Sharma, P.P. & Proud, D.N. (2016) Morphological tricks and blessed genitalia: rectifying the familial placement of Fijicolana tuberculata (Opiliones: Laniatores: Zalmoxidae). Zootaxa 4061:253-260. [PDF]
43. Clouse, R.M., Sharma, P.P., *Stuart, J.C., Davis, L.R., Giribet, G., Sarah, S.L. & Wheeler, W.C. (2016) Phylogeography of the harvestman Metasiro (Arthropoda, Arachnida, Opiliones) reveals a potential solution to the Pangean paradox. Organisms, Diversity and Evolution 16:167-184. [PDF] (*undergraduate co-author)
42. Sharma, P.P., *Santiago, M.A., González-Santillán, E., Monod, L. & Wheeler, W.C. (2015) Evidence of duplicated Hox genes in the most recent common ancestor of extant scorpions. Evolution and Development 17:347-355. [PDF] (*undergraduate co-author)